The Forum Herbulot world list
of family group names in Geometridae

The starting point of the "Forum Herbulot list of family group names of Geometridae" are the outstanding publications on the "Moths of Borneo" (Holloway 1994; 1996; 1997). The aim is to present all published names of the family group in Geometridae and to put them into a systematic order (without incorrect subsequent spellings). For this purpose a preliminary list has been discussed by the scientific board of the Forum Herbulot research initiative and is herewith presented on the homepage. The proposed systematic order and synonymous concepts need, of course, refinement and improvement by continuous discussion. The comittee thanks for personal contributions of Dr. E. Beljaev and Dr. A. Vives Moreno. Further proposals are highly welcome and may be addressed to the comittee. Systematic order and tribal concepts are main discussion focus at the Forum Herbulot congresses.

Available names in bold. Numbers of subordinated family group names in each head line of subfamilies.

The list is proposed to be cited as:
Forum Herbulot (2003): The Forum Herbulot world list of family group names in Geometridae <http://www.herbulot.de>, 11 pp., with pdf of update version of 12.6.2007 (A. Hausmann ed.)
Pdf of the world list, version 2 (12.6.2007)
Pdf of alternative order (Beljaev), version 2 (12.6.2007)

Familia Geometridae Stephens, 1829 (167)
Declared as correct name in Opinion 450.

Subfamilia Sterrhinae Meyrick, 1892 (Sterrhidae)(18)
Though there are several senior synonyms to this name (see below), the name should be maintained, in order to keep nomenclatural stability (as proposed by Holloway (1997: 15); see also Code art. 40.2. (ICZN 1999). Validity and order of tribes mainly basing on Holloway (1997) and Hausmann (2004).

Tribus Sterrhini Meyrick, 1892 (Sterrhidae)
Though there are senior synonyms to this name (see below), the name should be maintained, in order to keep nomenclatural stability (as proposed by Holloway (1997: 15) and Hausmann (2004: 20) ; see also Code art. 23.9 . (ICZN 1999).
=Goniacidaliini Packard, 1876 (Goniacidaliinae), senior synonym
=Idaeini Butler, 1881 (Idaeidae), senior synonym
=Ptychopodini Pierce, 1914 (Ptychopodinae), junior synonym

Tribus Scopulini Duponchel, 1845 (Scopulites)
=Acidaliini Duponchel, 1845 (Acidalites), emended to double ‚i‘, junior synonym, based on a junior homonym of a genus group name outside Lepidoptera
=Aletini Hampson, 1918 (Aletinae), junior subjective synonym (Holloway 1996)
=Problepsini Wiltshire, 1990 (Problepsini), junior subjective synonym (Holloway 1996)

Tribus Rhodostrophiini Prout, 1935 (Rhodostrophiicae)

Tribus Cyllopodini Kirby, 1892 (Cyllopodidae)
=Micropinini White, 1876 (Micropinidae), junior synonym, based on an unnecessary replacement genus-group name

Tribus Timandrini Stephens, 1850 (Timandridi)
=Calothysanini Herbulot, 1963 (Calothysanini), based on (misinterpreted) genus name Calothysanis, a synonym of Scopula (Scopulini).
Authorship: Agenjo, 1951 (Vives Moreno pers. comm. 2004)

Tribus Cosymbiini Prout, 1911 (Cosymbiinae)
Though there are senior synonyms to this name (see below), the name should be maintained, in order to keep nomenclatural stability (as proposed by Holloway (1997: 23); see also Code art. 40.2. (ICZN 1999).
=Ephyrini Guenée, 1858 (Ephyridae), based on a junior homonym of a genus group name outside Lepidoptera. Invalid according to Code art. 39 (ICZN 1999).
=Zonosomini White, 1876 (Zonosomatidi), basing on Zonosoma, thus emended, senior synonym
=Cyclophorini Moore, 1887 (Cyclophoridae), senior synonym; Cyclophoridae is in common usage in Mollusca based on the genus Cyclophorus, the name should be ‚gracefully conceded‘ to malacologists (Holloway 1997: 15)

Tribus Rhodometrini Agenjo, 1952 (Rhodometrinae)

Subfamilia Larentiinae Duponchel, 1845 (Larentites)(39)
Emended by reasons of grammar: basing on Larentia. Validity and order of tribes mainly basing on Holloway (1997) and Viidalepp (2007).

Tribus Lythriini Herbulot, 1962 (Lythriini)

Tribus Cataclysmini Herbulot, 1962 (Cataclysmini)

Tribus Xanthorhoini Pierce, 1914 (Xanthorhoinae)
=Euboliini Duponchel, 1845 (Eubolites), senior synonym (Holloway 1997: 97; Viidalepp 2007). Emended by reasons of grammar: basing on Eubolia, which is a junior synonym of Scotopteryx. It is proposed, not to apply this name, as Xanthorhoini is largely accepted (e.g. Holloway 1997, Viidalepp 2006), see Code art. 40.2 (ICZN 1999).
=Scotopterygini Warren, 1895 (Scotopteryginae), senior synonym (Holloway 1997: 98; Viidalepp 2006). Basing on Scotopteryx that falls within the concept of Xanthorhoini according to Viidalepp (2006). It is proposed, not to apply this name, as Xanthorhoini is largely accepted, see Code art. 23.9 (ICZN 1999).
=Ortholithini Tutt, 1896 (Ortholithinae), senior synonym (Holloway 1997: 98; Viidalepp 2007). Basing on Ortholitha, which is a junior synonym of Scotopteryx.
=Phasianini Gumppenberg, 1897 (Phasianinae). Basing on Phasiane, which is a junior synonym of Scotopteryx. Senior synonym, but junior homonym to a family group name in common use in Aves (Phasianus). (Holloway 1997: 98)
=Epirrhoini Pierce, 1914 (Epirrhoinae), junior synonym (Viidalepp 2007).
=Psychophorini Hampson, 1918 (Psychophorinae), junior synonym (Viidalepp 2006).

Tribus Euphyiini Herbulot, 1961 (Euphyiini)

Tribus Cidariini Duponchel, 1845 (Cidarites)
Emended by reasons of grammar: basing on Cidaria.
=Therini Pierce, 1914 (Therinae), junior synonym (Viidalepp 2006). Basing on Thera. Theriini (basing on Theria) is a family group name in Ennominae, basing on Theria.

Tribus Eudulini Warren, 1897 (Eudulinae)

Tribus Heterusiini Warren, 1897 (Heterusiinae)

Tribus Erateinini Guenée, 1858 (Erateinidae)
(Sometimes misspelled to Erateiniini).

Tribus Larentiini Duponchel, 1845 (Larentites)
Emended by reasons of grammar: basing on Larentia.
=Entephriini Pierce, 1914 (Entephrinae), emended by reasons of grammar (basing on Entephria), junior synonym (Viidalepp 2007).
=Mesoleucini McGuffin, 1958 (Mesoleucini), junior synonym (Viidalepp 2007).

Tribus Hydriomenini Meyrick, 1872 (Hydriomenidae)

Tribus Stamnodini Forbes, 1948 (Stamnodini)

Tribus Phileremini Pierce, 1914 (Philereminae)

Tribus Rheumapterini Herbulot, 1961 (Rheumapterini)
Authorship: Agenjo, 1951 (Vives Moreno pers. comm. 2004)
=Melanippini Bruand, 1846 (Melanippidae), senior synonym. Basing on Melanippe, which is a junior synonym of Rheumaptera. It is proposed, not to apply this name, as Rheumapterini is largely accepted.
=Eucosmiini Gumppenberg, 1887 (Eucosminae), senior synonym. [Applying of Code article 23.9.1.?] Emended by reasons of grammar (basing on Eucosmia, which is a junior synonym of Hydria). Eucosmini is a family group name in Tortricidae, basing on Eucosma.
=Triphosini Tutt, 1896 (Triphosidi), senior synonym, as falling in the concept of Rheumapterini (Viidalepp 2007).

Tribus Melanthiini Duponchel, 1845 (Melanthites)
Emended by reasons of grammar: basing on Melanthia.

Tribus Operophterini Packard, 1876 (Operophterinae)
=Oporiniini Pierce, 1914 (Oporiniinae), junior synonym (Holloway 1997: 98; Viidalepp 2007).
= Solitaneini Leraut, 1980 (Solitaneini). Nomen nudum and synonym (Vives Moreno pers. comm. 2004).

Tribus Asthenini Warren, 1894 (Astheninae)

Tribus Perizomini Herbulot, 1961 (Perizomini)

Tribus Eupitheciini Tutt, 1896 (Eupitheciinae)
Preferable over the prior name Tephroclystini according to Code art. 40.2. (ICZN 1999).
=Tephroclystiini Warren, 1895 (Tephroclystiinae), senior synonym. Based on Tephroclystia, a junior subjective synonym of Eupithecia.
=Chloroclystini Mironov, 1990 (Chloroclystina), junior synonym; proposed as subtribe of Eupitheciini.

Tribus Chesiadini Stephens, 1850 (Chesiadi)
Emended by reasons of grammar: basing on Chesias.
=Eucestini Warren, 1894 (Eucestiinae), emended by reasons of grammar (basing on Eucestis, which is a junior objective synonym of Chesias), junior synonym (Holloway 1997: 97).
=Odeziini Exposito, 1978 (Odeziini), junior synonym (Viidalepp 2007).

Tribus Trichopterygini Warren, 1894 (Trichopteryginae)
=Lobophorini Tutt, 1896 (Lobophorinae), junior synonym (Viidalepp: 2007).
= Dyspteridini Hulst, 1896: 249 (Dyspteridinae). Type genus Dyspteris Hübner, 1818. Dyspteridini need to be subordinated to Trichopterygini following the current treatment of genera composition of the latter. However there are morphological reasons for their separation (Beljaev pers. comm.).


Subfamilia Geometrinae Stephens, 1829 (Geometrinae)(29)
Validity and order of tribes mainly basing on Pitkin (1996) and Hausmann (1996a; 1996b; 2001).

Tribus Dysphaniini Warren, 1895 (Dysphaniinae)
=Hazini Guenée, 1858 (Hazidae), junior synonym; basing on Hazis, a junior objective synonym of Euschema, junior subjective synonym of Dysphania.
=Euschemini Walker, 1862 (Euschemidae), junior synonym; basing on Euschema, a junior synonym of Dysphania.

Tribus Pseudoterpnini Warren, 1893 (Pseudoterpninae)
=Terpnini Inoue, 1961 (Terpnini), junior synonym; based on Terpne (misspelled to Terpna), a junior synonym of Geometra, but referring to Terpne auctorum (=Pachyodes Guenée) (Hausmann, 1996a: 96; Holloway 1996)
=Archaeobalbini Viidalepp, 1981 (Archeobalbini), emended, junior synonym (Holloway 1996: 149)
=Pingasini Heppner & Inoue, 1992 (Pingasini), junior synonym

Tribus Heliotheini Exposito, 1978 (Heliothinae)
Emended, as basing on Heliothea. Subordinated under Geometrinae by Holloway (1996: 150) and Hausmann (1996b: 12; 2001: 110). Basal position in Geometrinae suggested in Hausmann (2001: 110); subordination under Rhomboristini suggested in Holloway (1996: 195)

Tribus Rhomboristini Inoue, 1961 (Rhomboristini)
Relationship to both Comostolini and Jodini suggested in Hausmann (1996b: 41).
=Lophochoristini Ferguson, 1969 (Lophochoristini), suggested as junior synonym (Holloway 1996: 195), according to Pitkin (1996: 322) valid at tribal rank.

Tribus Geometrini Stephens, 1829 (Geometrinae)
=Neohipparchini Inoue, 1961 (Neohipparchiti), perhaps junior synonym (compare Holloway 1996: 193)

Tribus Aracimini Inoue, 1961 (Aracimini)

Tribus Timandromorphini Inoue, 1961 (Timandromorphini)

Tribus Nemoriini Gumppenberg, 1887 (Nemorinae)
Emended by reasons of grammar.
=Ochrognesiini Inoue, 1961 (Ochrognesiini), junior synonym (Holloway 1996: 151; Hausmann 1996b: 31)

Tribus Dichordophorini Ferguson, 1969 (Dichordophorini)

Tribus Synchlorini Ferguson, 1969 (Synchlorini)
Relations to Comibaenini discussed in Holloway (1996) and Pitkin (1996: 322).

Tribus Comibaenini Inoue, 1961 (Comibaenini)
=Euchlorini Herbulot, 1963 (Euchlorini), junior synonym; based on genus name Euchloris, which is a junior homonym of a generic name in Coleoptera.
=Synchlorini Ferguson, 1969 (Synchlorini), junior synonym (Holloway 1996: 196)

Tribus Hemistolini Inoue, 1961 (Hemistolini)
For validity of this name at tribe rank see Hausmann (1996b; 2001), Holloway (1996: 196) proposes a wider concept (Thalerini, Comostolini, Hemistolini, Jodini and Thalassodini joined to one single (sub)tribe).

Tribus Thalassodini Inoue, 1961 (Thalassodini)
For validity of this name at tribe rank see Hausmann (1996b), Holloway (1996: 196) proposes a wider concept (Thalerini, Comostolini, Hemistolini, Jodini and Thalassodini joined to one single (sub)tribe).

Tribus Jodini Inoue, 1961 (Jodiini)
Emended by reasons of grammar (basing on Jodis). For validity of this name at tribe rank see Hausmann (1996b; 2001), related to Comostolini (Hausmann 1996b: 41). Holloway (1996: 196) proposes a wider concept (Thalerini, Comostolini, Hemistolini, Jodini and Thalassodini joined to one single (sub)tribe).

Tribus Comostolini Inoue, 1961 (Comostolini)
For validity of this name at tribe rank see Hausmann (1996b; 2001), related to Jodini (Hausmann 1996b: 41). Holloway (1996: 196) proposes a wider concept (Thalerini, Comostolini, Hemistolini, Jodini and Thalassodini joined to one single (sub)tribe).

Tribus Thalerini Herbulot, 1963 (Thalerini)
For validity of this name at tribe rank see Hausmann (1996b; 2001), Holloway (1996: 196) proposes a wider concept (Thalerini, Comostolini, Hemistolini, Jodini and Thalassodini joined to one single (sub)tribe).
=Chlorochromini Duponchel, 1845 (Chlorochromites), senior synonym, based on Chlorochroma, a junior objective synonym of Thalera. It is proposed, not to apply this name, as Thalerini is largely accepted (see Hausmann 1996a: 101).

Tribus Hemitheini Bruand, 1846 (Hemitheidae)
For validity of this name at tribe rank see Hausmann (1996; 2001), Holloway (1996: 196) proposes a wider concept (Thalerini, Comostolini, Hemistolini, Jodini and Thalassodini joined to one single (sub)tribe).

Tribus Microloxiini Hausmann, 1996 (Microloxiini)
Published on 21.xi.
=Hierochthoniini Viidalepp, 1996, published without exact date (=31.xii.)

Subfamilia Archiearinae Fletcher, 1953 (Archiearinae)(2)
=Brephinae auct. nec Hübner, 1826 (Brephidae), basing on Brephos Ochsenheimer, 1816, a junior homonym of Brephos Hübner, 1813.

Subfamilia Oenochrominae Guenée, 1858 (Oenochromidae)(3)
=Lyrceini Meyrick, 1883 (Lyrceini), basing on Lyrcea, a junior homonym of a name outside Lepidoptera; junior subjective synonym. Invalid according to Code art. 39 (ICZN 1999).
=Monocteniini Meyrick, 1889 (Monocteniadae), emended, junior subjective synonym (Holloway 1996: 151)

Subfamilia Desmobathrinae Meyrick, 1886 (Desmobathridae)(4)
Based on Desmobathra (junior synonym of Ozola). For validity at subfamily rank see Holloway (1996: 152, 159)
=Mecoceratini Guenée, 1858 (Mecoceridae), emended by Hulst (1896) by reasons of grammar, basing on Mecoceras. The latter generic name being an objective synonym of Ametris.
=Ametridini Prout, 1910 (Ametridicae), junior synonym (synonymy to be confirmed)

Tribus Desmobathrini Meyrick, 1886 (Desmobathridae)
=Epirrhanthini Forbes, 1948 (Epirrhanthini), junior synonym (synonymy to be confirmed) and subordinated here in Hausmann (2001: 99), under Ennominae in Holloway 1994: 7).

Tribus Eumeleini Warren, 1894 (Eumeleinae)

Subfamilia Orthostixinae Meyrick, 1892 (Orthostixidae)(1)
Subfamily rank questioned (Holloway 1996: 151), possibly subordinated to Ennominae; validity at subfamily rank suggested by Hausmann (2001: 70, 89)

Subfamilia Ennominae Duponchel, 1845 (Ennomites)(72)
Validity and order of tribes mainly basing on Holloway (1994) and Pitkin (2002).

See also alternative order communicated by E. Beljaev (2006)

Tribus Diptychini Janse, 1933 (Diptychini)
Subfamily rank and position close to Oenochrominae/Desmobathrinae has repeatedly been suggested. For subordination under Ennominae see Holloway (1996: 150). Relationships to Nacophorini and Ourapterygini see Holloway (1996: 151) and Pitkin (2002: 135).

Tribus Nacophorini Forbes, 1948 (Nacophorini)
Relationship to Azelini suggested by Holloway (1994: 8)

Tribus Azelinini Forbes, 1948 (Azelinini)
Relationship to Nacophorini suggested by Holloway (1994: 5)

Tribus Odontoperini Tutt, 1896 (Odontoperinae)
See remarks in Holloway (1994: 8, 111). Related to Nacophorini/Azelini (Abraham et al. 2001: fig. 5b).

Tribus Hypochrosini Guenée, 1858 (Hypochrosinae)
=Anagogini Forbes, 1948 (Anagogini), junior subjective synonym (Holloway: 1994: 5).

Tribus Scardamiini Warren, 1894 (Scardamiinae)

Tribus Ourapterygini Bruand, 1846 (Urapteridae)
Emended by Forbes (1848), Holloway (1996: 6).
=Emplociini Guenée, 1858 (Emplocidae). Emended by reasons of grammar. Junior subjective synonym (Pitkin 2002: 135).
=Oxydiini Butler, 1886 (Oxydiidae). Junior subjective synonym (Pitkin 2002: 135).
=Cingiliini Forbes, 1948 (Cingiliini). Junior subjective synonym (Holloway 1994: 6; Pitkin 2002: 135).

Tribus Epionini Bruand, 1846 (Epionidae)
=Rumiini Tutt, 1896 (Rumiinae), basing on Rumia, which is a junior objective synonym of Opisthograptis. Junior subjective synonym (Leraut 1997).

Tribus Angeronini Forbes, 1948 (Angeronini)
=Aspilatini Duponchel, 1845 (Aspilatites), senior subjective synonym (Holloway: 1994: 5), but basing on a misspelled genus group name (Aspitates) (Holloway: 1994: 5).
Emendation: Aspitatini Leraut, 1997

Tribus Ennomini Duponchel, 1845 (Ennomites)
=Odopterini Stephens, 1850 (Odopteridi), basing on Odoptera, an unnecessary replacement name of Ennomos.
=Crocallini Tutt, 1896 (Crocallidi), proposed here as junior subjective synonym.
=Seleniini Tutt, 1896 (Seleniidi), suggested as junior subjective synonym in Holloway (1994: 9).

Tribus Nephodiini Warren, 1894 (Nephodiinae)

Tribus Plutodini Warren, 1894 (Plutodinae)

Tribus Colotoini Wehrli, 1940 (Colotoinae)

Tribus Thinopterygini Holloway, 1994 (Thinopterygini)

Tribus Gonodontini Forbes, 1948 (Gonodontini)
See remarks in Holloway (1994: 8, 111).

Tribus Abraxini Warren, 1893 (Abraxinae)
=Zerenini Duponchel, 1845 (Zerenites), based on a junior homonym of a genus group name outside Lepidoptera; junior subjective synonym (Holloway 1994: 9). Invalid according to Code art. 39 (ICZN 1999).
=Pantherini Moore, 1887 (Pantheridae); based on a junior homonym of a genus group name outside Lepidoptera (Holloway 1994: 9); invalid according to Code art. 39 (ICZN 1999). Tentatively placed here as junior subjective synonym.

Tribus Cassymini Holloway, 1994 (Cassymini)

Tribus Eutoeini Holloway, 1994 (Eutoeini)

Tribus Macariini Guenée, 1858 (Macaridae)
Emended by reasons of grammar (basing on Macaria).
=Semiothisini Warren, 1894 (Semiothisinae), junior synonym (Holloway 1994: 9).
=Fernaldellini Hulst, 1896 (Fernaldellinae), junior subjective synonym (Holloway 1994: 7).

Tribus Boarmiini Duponchel, 1845 (Boarmites)
A very broad conception of this tribe has been proposed by Holloway (1994: 167)
Strict concept:
=Cleorini Duponchel, 1845 (Cleorites), junior subjective synonym (Holloway: 1994: 6).
=Fidoniini Duponchel, 1845 (Fidonites), emended by reasons of grammar; basing on Fidonia, a junior synonym of Eurranthis, junior subjective synonym (Leraut 1997: 214).
=Ascotini Warren, 1893 (Ascotinae), junior subjective synonym (Holloway: 1994: 5).
‚Subgroup‘Bistonini
=Bistonini Stephens, 1850 (Bistonidi)
Falling into thebroad conception of Boarmiini according to Holloway (1994: 13, 167)
=Amphidasini Duponchel, 1845 (Amphidasites), senior synonym. Basing on Amphidasis, which is a junior objective synonym of Biston. It is proposed, not to apply this name, as Bistonini is largely accepted (see Holloway 1994: 5).
=Hyberniini Duponchel, 1845 (Hibernites), senior synonym. Emended. Basing on Hybernia, which is a junior objective synonym of Erannis. It is proposed, not to apply this name, as Bistonini is largely accepted (see Holloway 1994: 5,7).
=Eubyjini Warren, 1893 (Eubyjinae), basing on Eubyja, which is a junior subjective synonym of Biston (Holloway 1994: 7).
=Erannini Tutt, 1896 (Eranniinae), junior synonym (Holloway 1994: 7). Emended.
Wide concept:
=Dalimini Wehrli, 1940 (Daliminae), junior subjective synonym (Holloway 1996: 150).
=Selidosemini Meyrick, 1892 (Selidosemidae), junior subjective synonym (Holloway 1994: 9).
=Braccini Warren, 1894 (Braccinae), junior subjective synonym (Holloway 1994: 167).
=Melanchroiini Hulst, 1896 (Melanchroiinae), junior subjective synonym (Pitkin 2002: 130).
=Phaseliini Wehrli, 1941 (Phaseliinae), suggested to fall into the wide concept of Boarmiini.
=Melanolophini Forbes, 1948 (Melanolophini), junior subjective synonym (Holloway 1994: 8; Pitkin 2002: 131).
=Bupalini Herbulot, 1963 (Bupalini), junior subjective synonym (Holloway 1994: 6; 167).
=
Milioniini Holloway, 1994 (Milioniini). First mentioned by Inoue, 1992, but without description, unavailable. Junior subjective synonym (Holloway 1994: 8, 302).
=Obeidiini Holloway, 1994 (Obeidiini). First mentioned by Inoue, 1992, but without description, unavailable. Junior subjective synonym (Holloway 1994: 8, 302).

Tribus Theriini Herbulot, 1963 (Theriini)
Falling into thebroad conception of Boarmiini according to Holloway (1994: 13, 167)
=Cheimatobiini Tutt, 1896 (Cheimatobiidi), senior synonym. Basing on Cheimatobia, which is a junior objective synonym of Theria. It is proposed, not to apply this name, as Theriini is largely accepted (see Holloway 1994: 6).

Tribus Gnophini Duponchel, 1845 (Gnophites)
Suggested to fall, probably, under the wide concept of Boarmiini by Holloway (1994: 7)
=Dasydini Duponchel, 1845 (Dasydites), basing on Dasydia, a junior objective synonym of Sciadia. Possibly valid at tribe rank (Erlacher pers. comm.).
=Sionini Duponchel, 1845 (Sionites), junior subjective synonym (see Holloway 1994: 9). Possibly valid at tribe rank (Erlacher pers. comm.).

Tribus Campaeini Forbes, 1948 (Campaeini)
=Metrocampini Tutt, 1896 (Metrocampidae), senior synonym. Basing on Metrocampa, which is a junior objective synonym of Campaea. It is proposed, not to apply this name, as Campaeini is largely accepted (see Holloway 1994: 6).

Tribus Alsophilini Herbulot, 1962 (Alsophilinae)
Downgraded from subfamily to tribe rank after getting the same result in several nDNA studies (e.g. Young 2006; Yamamoto & Sota 2007). This group seems to be closely related with Campaeini and Colotoini.

Tribus Lithinini Forbes, 1948 (Lithinini)
Closely related to Caberini (Abraham et al. 2001: fig. 5b).
=Lacariini Orfila & Schajovskoy, 1959 (Lacarini), emended. For possible synonymy to Lithinini see Holloway (1994: 7, 92)

Tribus Caberini Duponchel, 1845 (Caberites)
Closely related to Lithinini (Abraham et al. 2001: fig. 5b).
=Erastriini Herrich-Schäffer, 1845 (Erastridae), junior synonym (Holloway 1996: 7; 98). Emended. The original concept referring to acontiine Noctuids, Because of the misidentification of the generic type species the name is now applicable within Ennominae (Holloway 1994: 7).
=Brotini Grote, 1882 (Brotiinae), Emended. Basing on Brotis, which is a junior homonym to a genus group name in Noctuidae. Invalid according to Code art. 39 (ICZN 1999). Junior synonym in the broad concept of Caberini/Baptini of Pitkin (2002: 131).
=Deiliniini Warren, 1894 (Deiliniinae), basing on Deilinia, a junior synonym of Cabera (Holloway 1996: 6; 98).
=Catopyrrhini Warren, 1894 (Catopyrrhinae), junior subjective synonym (Holloway 1996: 6; 98).
=Sphacelodini Forbes, 1948 (Sphacelodini), retained synonym in the broad concept of Caberini/Baptini of Pitkin (2002: 131).

- (Sub-)Tribus Baptini Forbes, 1948 (Baptini)
Closely related to Palyadini (Abraham et al. 2001: fig. 5b), suggested to be subordinated under Caberini in Pitkin (2002: 131).

- (Sub-)Tribus Palyadini Guenée, 1858 (Palyadae)
Emended by reasons of grammar. Subordination under Baptini rejected by Holloway (1994: 59), but closely related to Baptini (Abraham et al. 2001: fig. 5b). Pitkin (2002: 132) suggests status as subtribe of Caberini/Baptini.

Tribus Prosopolophini Warren, 1894 ( Prosopolophinae )
= Pachycnemiini Kirby, 1903 (Pachycnemiidae) junior synonym (Beljaev, Vives Moreno, pers. comm. 2004)
=Ligiini Guenée, 1858 (Ligidae), emended. Senior synonym, but basing on a junior homonym of a genus group name outside Lepidoptera. Invalid according to Code art. 39 (ICZN 1999).
= Compsopterini Herbulot, 1963 (Compsopterini) junior synonym (Beljaev, Vives Moreno, pers. comm. 2004)

Tribus Onychorini Herbulot, 1963 (Onychorini)


References:

Abraham, D., Ryrholm, N., Wittzell, H., Holloway, J.D., Scoble, M.J. & C. Löfstedt 2001. Molecular Phylogeny of the Subfamilies in Geometridae (Geometroidea: Lepidoptera). – Mol. Phylo. Evol. 20 (1): 65-77

Beljaev, E. A. 1994. [Geometers of the subfamily Ennominae (Lepidoptera, Geometridae) of the Far East of Russia. Abstract of the candidate of biological sciences thesis. ] 23 pp. Zoological institute of Russian Academy of Sciences. St.-Petersburg. (In Russian.)

Beljaev, E. A. 1997. A new genus and new species of the Ennominae (Lepidoptera, Geometridae) from Russian Far East, with remarks on the phylogeny of the related genera. Tinea 15(2): 125-130.

Beljaev, E. A. 1998. Morphology of the copulatory apparatus in Devenilia corearia and description of a new tribe Deveniliini (Lepidoptera, Geometridae, Ennominae). Zoologicheskii Zhurnal 77(4): 438-443.

Beljaev, E. A. 2000. Remarkable new genus and new species of the geometrid moths from Central Asia , related to the genus Desertobia Viidallepp, 1989 (Lepidoptera, Geometridae, Ennominae) with notes on the taxonomy of the Desertobiini. Tinea 16 (4): 240-245.

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